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Elephant seals are known to engage in bloody battles in order to retain their title has dominant male so that they are able to mate with all the females in their community.

Copulatory plugs are frequently observed in insects, reptiles, some mammals, and spiders. This behavior ensures males with higher mating success at the expense of female control of sperm sperm selection.

Similarly, Drosophila melanogaster males release toxic seminal fluids, known as ACPs accessory gland proteins , from their accessory glands to impede the female from participating in future copulations.

Another strategy, known as sperm partitioning, occurs when males conserve their limited supply of sperm by reducing the quantity of sperm ejected.

A strategy common among insects is for males to participate in prolonged copulations. By engaging in prolonged copulations a male has an increased opportunity to place more sperm within the female's reproductive tract and prevent the female from copulating with other males.

It has been found that some male mollies Poecilia have developed deceptive social cues to combat sperm competition. Focal males will direct sexual attention toward typically non-preferred females when an audience of other males is present.

This encourages the males that are watching to attempt to mate with the non-preferred female. This is done in an attempt to decrease mating attempts with the female that the focal male prefers, hence decreasing sperm competition.

Offensive adaptation behavior differs from defensive behavior because it involves an attempt to ruin the chances of another male's opportunity in succeeding in copulation by engaging in an act that tries to terminate the fertilization success of the previous male.

Further, the presence of an offensive trait sometimes serves as a status signal. The mere display of an armament can suffice to drive away the competition without engaging in a fight, hence saving energy.

The "good sperm hypothesis" is very common in polyandrous mating systems. Studies show that there is more to determining the competitiveness of the sperm in sperm competition in addition to a male's genetic makeup.

A male's dietary intake will also affect sperm competition. An adequate diet consisting of increased amounts of diet and sometimes more specific ratio in certain species will optimize sperm number and fertility.

Amounts of protein and carbohydrate intake were tested for its effects on sperm production and quality in adult fruit flies Diptera: Tephritidae.

Studies showed these flies need to constantly ingest carbohydrates and water to survive, but protein is also required to attain sexual maturity.

In addition, protein and carbohydrate amounts were shown to have an effect on sperm production and fertility in the speckled cockroach. Holidic diets were used which allowed for specific protein and carbohydrate measurements to be taken, giving it credibility.

A direct correlation was seen in sperm number and overall of food intake. More specifically, optimal sperm production was measured at a protein to carbohydrate ratio.

Sperm fertility was best at a similar protein to carbohydrate ratio of This close alignment largely factors in determining male fertility in Nauphoeta cinerea.

It's hypothesized that sperm viability is more affected by the genetic makeup, like in the "good sperm hypothesis". These ratios and results are not consistent with many other species and even conflict with some.

It seems there can't be any conclusions on what type of diet is needed to positively influence sperm competition but rather understand that different diets do play a role in determining sperm competition in mate choice.

One evolutionary response to sperm competition is the variety in penis morphology of many species. Evolution to increase ejaculate volume in the presence of sperm competition has a consequence on testis size.

Large testes can produce more sperm required for larger ejaculates, and can be found across the animal kingdom when sperm competition occurs.

In some insects and spiders, for instance Nephila fenestrate , the male copulatory organ breaks off or tears off at the end of copulation and remains within the female to serve as a copulatory plug.

Female factors can influence the result of sperm competition through a process known as "sperm choice". One instance where this is known to occur is inbreeding; females will preferentially use the sperm from a more distantly related male than a close relative.

Inbreeding ordinarily has negative fitness consequences inbreeding depression , and as a result species have evolved mechanisms to avoid inbreeding.

Inbreeding depression is considered to be due largely to the expression of homozygous deleterious recessive mutations.

Numerous inbreeding avoidance mechanisms operating prior to mating have been described. However, inbreeding avoidance mechanisms that operate subsequent to copulation are less well known.

In guppies , a post-copulatory mechanism of inbreeding avoidance occurs based on competition between sperm of rival males for achieving fertilization.

In vitro fertilization experiments in the mouse, provided evidence of sperm selection at the gametic level. The results were interpreted as egg-driven sperm selection against related sperm.

Female fruit flies Drosophila melanogaster were mated with males of four different degrees of genetic relatedness in competition experiments.

Female crickets Teleogryllus oceanicus appear to use post-copulatory mechanisms to avoid producing inbred offspring.

When mated to both a sibling and an unrelated male, females bias paternity towards the unrelated male. It has been found that because of female choice see sexual selection , morphology of sperm in many species occurs in many variations to accommodate or combat see sexual conflict the morphology and physiology of the female reproductive tract.

Polyandrous females mate with many male partners. Males will invest more in ejaculation when competitors are in the immediate environment of the female.

Evidence exists that illustrates the ability of genetically similar spermatozoa to cooperate so as to ensure the survival of their counterparts thereby ensuring the implementation of their genotypes towards fertilization.

Cooperation confers a competitive advantage by several means, some of these include incapacitation of other competing sperm and aggregation of genetically similar spermatozoa into structures that promote effective navigation of the female reproductive tract and hence improve fertilization ability.

Such characteristics lead to morphological adaptations that suit the purposes of cooperative methods during competition. For example, spermatozoa possessed by the wood mouse Apodemus sylvaticus possess an apical hook which is used to attach to other spermatozoa to form mobile trains that enhance motility through the female reproductive tract.

Other evidence suggests no link between sperm competition and sperm hook morphology. Selection to produce more sperm can also select for the evolution of larger testes.

Relationships across species between the frequency of multiple mating by females and male testis size are well documented across many groups of animals.

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